Penelopides

Penelopides
Visayan hornbill (Penelopides panini)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Bucerotiformes
Family:	Bucerotidae
Genus:	Penelopides Reichenbach, 1849
Type species
Buceros panini[1] Boddaert, 1783
Species
2-5, see text

Penelopides is the genus of relatively small, primarily frugivorous hornbills restricted to forested areas of the Philippines. Their common name, tarictic hornbills, is an onomatopoetic reference to the main call of several of them. They have a ridged plate-like structure on the base of their mandible. All are sexually dimorphic: males of all species are whitish-buff and black, while females of all except the Mindoro hornbill are primarily black.

Taxonomy

The genus Penelopides was introduced in 1849 by the German naturalist Ludwig Reichenbach in a plate of the hornbills.^[2] The type species was subsequently designated as the Visayan hornbill (Penelopides panini) by the English zoologist George Gray.^[3][4] The origin of Reichenbach's generic name is uncertain. It may be a combination of the Latin pene meaning "almost" or "nearly", the Ancient Greek lophos meaning "crest" and -oidēs "resembling".^[5]

While the taxonomic history of the Sulawesi hornbill has been uneventful, the number of species in the Philippine Archipelago has been greatly debated. Traditionally, only a single Philippine species was recognized, the tarictic hornbill (P. panini). Following a review, it was recommended that it should be split into four allopatric species, the Visayan (P. panini), Luzon (P. manillae), Mindanao (P. affinis), and Mindoro (P. mindorensis). Today most authorities accept this split, although some have recommended splitting the Samar (P. samarensis) from the Mindanao.^{[clarification needed]}

The genus contains five species:^[6]

Image	Scientific name	Common Name	Distribution
	Penelopides panini	Visayan hornbill	islands of Panay, Negros, Masbate, and Guimaras, and formerly Ticao, in the Philippines.
	Penelopides manillae	Luzon hornbill	Luzon and nearby islands in the northern Philippines
	Penelopides affinis	Mindanao hornbill	Mindanao, Dinagat, Siargao and Basilan in the southern Philippines
	Penelopides samarensis	Samar hornbill	islands of Samar, Calicoan, Leyte and Bohol in the east-central Philippines
	Penelopides mindorensis	Mindoro hornbill	Mindoro in the Philippines

References

• Kemp, A. C. (2001). Family Bucerotidae (Hornbills). pp. 436–523 in: del Hoyo, J., Elliott, A., & Sargatal, J. eds. (2001). Handbook of the Birds of the World. Vol. 6. Mousebirds to Hornbills. Lynx Edicions, Barcelona. ISBN 84-87334-30-X

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Genera of trogons, hornbills, kingfishers, woodpeckers and their extinct allies

• Kingdom: Animalia
• Phylum: Chordata
• Class: Aves
• Infraclass: Neognathae
• Clade: Coraciimorphae
• Clade: Cavitaves

Cavitaves
Leptosomiformes Leptosomidae Leptosomus †Plesiocathartes? †Waltonavis Eucavitaves Trogoniformes Trogonidae Apalharpactes Apaloderma †Eotrogon Euptilotis †Foshanornis Harpactes †Masillatrogon †Paratrogon Pharomachrus †Primotrogon Priotelus †Septentrogon Trogon Picocoraciae Bucerotiformes See below ↓ Picodynastornithes Coraciiformes See below ↓ Piciformes See below ↓

Bucerotiformes
Bucerotiformes †Laurillardiidae Laurillardia †Messelirrisoridae Messelirrisor Upupi Phoeniculidae †Phirriculus Phoeniculus Rhinopomastus Upupidae Upupa Buceroti Bucorvidae Bucorvus †Euroceros Bucerotidae Aceros Anorrhinus Anthracoceros Berenicornis Buceros Bycanistes Ceratogymna Horizocerus Lophoceros Ocyceros Penelopides Rhabdotorrhinus Rhinoplax Rhyticeros Tockus

Coraciiformes
incertae sedis †Cryptornis †Paracoracias †Quasisyndactylus †Eocoraciidae Eocoracias †Geranopteridae Geranopterus †Primobucconidae Primobucco Septencoracias Brachypteraciidae Atelornis Brachypteracias Geobiastes Uratelornis Coraciidae †Chehuenia Coracias Eurystomus †Miocoracias Meropidae Meropogon Merops Nyctyornis Alcedines Alcedinidae Alcedininae Alcedo Ceyx Corythornis Ispidina Cerylinae Ceryle Chloroceryle Megaceryle Halcyoninae Actenoides Caridonax Cittura Clytoceyx Dacelo Halcyon Lacedo Melidora Pelargopsis Syma Tanysiptera Todiramphus Momotidae Aspatha Baryphthengus Electron Eumomota Hylomanes Momotus †Protornis? Todidae †Palaeotodus Todus

Piciformes
incertae sedis †"Homalopus" †Gracilitarsidae Gracilitarsus Neanis †Sylphornithidae Oligosylphe Palaegithalus? Sylphornis Galbuli †Jacamatia Bucconidae Bucco Chelidoptera Hapaloptila Hypnelus Malacoptila Micromonacha Monasa Nonnula Notharchus Nystalus Galbulidae Brachygalba Galbalcyrhynchus Galbula Jacamaralcyon Jacamerops Pici incertae sedis †Rupelramphastoides †Miopiconidae Miopico †Picavidae Picavus Ramphastides Capitonidae Capito †Capitonides Eubucco Lybiidae Buccanodon Cryptolybia Gymnobucco Lybius Pogoniulus Pogonornis Stactolaema Trachyphonus Tricholaema Megalaimidae Caloramphus Psilopogon Ramphastidae Andigena Aulacorhynchus Pteroglossus Ramphastos Selenidera Semnornithidae Semnornis Picides Indicatoridae Indicator Melichneutes Melignomon Prodotiscus Picidae See below ↓

Picidae
incertae sedis †Bitumenpicus †Breacopus †Palaeonerpes †Palaeopicus †Piculoides Jynginae Jynx Picumninae Picumnus Sasiinae Sasia Verreauxia Picinae †Australopicus †Palaeonerpes †Pliopicus Nesoctitini Nesoctites Hemicircini Hemicircus Picini Campethera Celeus Chrysophlegma Colaptes Dinopium Dryocopus Gecinulus Geocolaptes Micropternus Meiglyptes Mulleripicus Pardipicus Piculus Picus Campephilini Blythipicus Campephilus Chrysocolaptes Reinwardtipicus Melanerpini Chloropicus Dendrocopos Dendrocoptes Dendropicos Dryobates Leiopicus Leuconotopicus Melanerpes Picoides Sphyrapicus Veniliornis Xiphidiopicus Yungipicus

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