Suborder of beetles
Archostemata
Temporal range: Late Permian–Recent PreꞒ Ꞓ O S D C P T J K Pg N |
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| Tenomerga mucida |
Scientific classification  |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | Archostemata
Kolbe, 1908 |
| Families |
| Crowsoniellidae
Cupedidae
Jurodidae
Micromalthidae
Ommatidae |
The Archostemata are the smallest suborder of beetles, consisting 45 living species in five families.[1] They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, Micromalthus debilis. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like "protocoleopterans", which some modern archostematans closely resemble to due to their plesiomorphic morphology.[2] Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga.[1]
Taxonomy
There are five extant families.[3][4]
See also
References
- ^ a b Hörnschemeyer, Thomas (July 2009). "The species-level phylogeny of archostematan beetles—where do Micromalthus debilis and Crowsoniella relicta belong?". Systematic Entomology. 34 (3): 533–558. doi:10.1111/j.1365-3113.2009.00476.x. S2CID 84795808.
- ^ Boudinot, Brendon Elias; Yan, Evgeny Viktorovich; Prokop, Jakub; Luo, Xiao‐Zhu; Beutel, Rolf Georg (2022-07-28). "Permian parallelisms: Reanalysis of †Tshekardocoleidae sheds light on the earliest evolution of the Coleoptera". Systematic Entomology. 48: 69–96. doi:10.1111/syen.12562. ISSN 0307-6970. S2CID 251171914.
- ^ Patrice Bouchard; Yves Bousquet; Anthony E. Davies; Miguel A. Alonso-Zarazaga; John F. Lawrence; Chris H. C. Lyal; Alfred F. Newton; Chris A. M. Reid; Michael Schmitt; S. Adam Ślipiński; Andrew B. T. Smith (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
- ^ Bouchard, P.; Bousquet, Y. (2020). "Additions and corrections to "Family-group names in Coleoptera (Insecta)"". ZooKeys (922): 65–139. doi:10.3897/zookeys.922.46367. PMC 7113323. PMID 32256157.
External links
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Suborder Archostemata |
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- Crowsoniellidae (Crowsoniella relicta)
- Cupedidae (reticulated beetles)
- Jurodidae (Sikhotealinia zhiltzovae)
- Micromalthidae (telephone-pole beetle)
- Ommatidae
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| Extant families | - Amphizoidae (trout-stream beetles)
- Aspidytidae
- Carabidae (ground beetles)
- Cicindelidae (tiger beetles)
- Dytiscidae (predaceous diving beetles)
- Gyrinidae (whirligig beetles)
- Haliplidae (crawling water beetles)
- Hygrobiidae
- Meruidae (Meru phyllisae)
- Noteridae (burrowing water beetles)
- Trachypachidae (false ground beetles)
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| Bostrichiformia | | Bostrichoidea | - Bostrichidae (auger beetles)
- Dermestidae (skin beetles)
- Endecatomidae
- Jacobsoniidae (Jacobson's beetles)
- Nosodendridae (wounded-tree beetles)
- Ptiniidae (furniture beetles, death watch beetles, spider beetles)
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| Derodontoidea | - Derodontidae (tooth-necked fungus beetles)
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| Cucujiformia | | Chrysomeloidea | |
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| Cleroidea | |
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| Coccinelloidea | |
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| Cucujoidea | |
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Curculionoidea
(weevils) | - Anthribidae (fungus weevils)
- Attelabidae (leaf-rolling weevils)
- Belidae (primitive weevils)
- Brentidae (straight snout weevils, New York weevil)
- Caridae
- Curculionidae (true weevils, bark beetles, ambrosia beetles)
- Nemonychidae (pine flower weevils)
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| Lymexyloidea | - Lymexylidae (ship-timber beetles)
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| Tenebrionoidea | - Aderidae (ant-like leaf beetles)
- Anthicidae (ant-like flower beetles)
- Archeocrypticidae (cryptic fungus beetles)
- Boridae (conifer bark beetles)
- Chalcodryidae
- Ciidae (minute tree-fungus beetles)
- Melandryidae (false darkling beetles)
- Meloidae (blister beetles)
- Mordellidae (tumbling flower beetles)
- Mycetophagidae (hairy fungus beetles)
- Mycteridae (palm and flower beetles)
- Oedemeridae (false blister beetle)
- Perimylopidae, or Promecheilidae
- Prostomidae (jugular-horned beetles)
- Pterogeniidae
- Pyrochroidae (fire-coloured beetles)
- Pythidae (dead log bark beetles)
- Ripiphoridae (wedge-shaped beetles)
- Salpingidae (narrow-waisted bark beetles)
- Scraptiidae (false flower beetles)
- Stenotrachelidae (false longhorn beetles)
- Synchroidae (synchroa bark beetles)
- Tenebrionidae (darkling beetles)
- Tetratomidae (polypore fungus beetles)
- Trictenotomidae
- Ulodidae
- Zopheridae (ironclad beetles, cylindrical bark beetles)
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| Elateriformia | | Buprestoidea | - Buprestidae (jewel beetles, or metallic wood-boring beetles)
- Schizopodidae
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| Byrrhoidea | - Byrrhidae (pill beetles)
- Callirhipidae (cedar beetles)
- Chelonariidae (turtle beetles)
- Cneoglossidae
- Dryopidae (long-toed water beetles)
- Elmidae (riffle beetles)
- Eulichadidae (forest stream beetles)
- Heteroceridae (variegated mud-loving beetles)
- Limnichidae (minute mud beetles)
- Lutrochidae (travertine beetles)
- Psephenidae (water-penny beetles)
- Ptilodactylidae
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| Dascilloidea | - Dascillidae (soft bodied plant beetles)
- Rhipiceridae (cicada beetle, cicada parasite beetles)
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| Elateroidea | - Artematopodidae (soft-bodied plant beetles)
- Brachypsectridae (Texas beetles)
- Cantharidae (soldier beetles)
- Cerophytidae (rare click beetles)
- Elateridae (click beetles)
- Eucnemidae (false click beetles)
- Jurasaidae
- Lampyridae (fireflies)
- Lycidae (net-winged beetles)
- Omethidae (false fireflies, long-lipped beetles)
- Phengodidae (glowworm beetles)
- Rhagophthalmidae
- Sinopyrophoridae
- Throscidae (false metallic wood-boring beetles)
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| Rhinorhipoidea | - Rhinorhipidae (Rhinorhipus tamborinensis)
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| Scirtoidea | |
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| Scarabaeiformia | | Scarabaeoidea | - Belohinidae (Belohina inexpectata)
- Bolboceratidae
- Diphyllostomatidae (false stag beetles)
- Geotrupidae (dor beetles)
- Glaphyridae (bumble bee scarab beetles)
- Glaresidae (enigmatic scarab beetles)
- Hybosoridae (scavenger scarab beetles)
- Lucanidae (stag beetles)
- Ochodaeidae (sand-loving scarab beetles)
- Passalidae (betsy beetles)
- Pleocomidae (rain beetles)
- Scarabaeidae (scarabs)
- Trogidae (hide beetles)
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| Staphyliniformia | |
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